Multiple lines of evidence have further demonstrated that PRRs directly bind the promoters of PHYTOCHROME-INTERACTING FACTOR4 ( PIF4) and PIF5 to repress their expression, hence PRRs act as transcriptional repressors of the positive growth regulators PIF4 and PIF5. Intriguingly, a distinct daylength could shift the expression phase and extend the expression duration of PRRs. Here, we show that Pseudo Response Regulators (PRRs) coordinate with the Evening Complex (EC), a transcriptional repressor complex within the clock core oscillator, to specifically regulate photoperiodic hypocotyl growth in Arabidopsis ( Arabidopsis thaliana). The circadian clock measures and conveys daylength information to control rhythmic hypocotyl growth in photoperiodic conditions to achieve optimal fitness, but it operates through largely unknown mechanisms.
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